Accumulation of constitutive diterpenoids in the rhytidome and secondary phloem of the branch bark of Larix gmelinii var. japonica

Journal of Wood Science

Official Journal of the Japan Wood Research Society

Table 1 Contents of the petroleum ether–diethyl ether extract and composition of the constitutive diterpenoids in the rhytidome and secondary phloem in the branch bark of Larix gmelinii var. japonica

	Rhytidome (mg/g)^a			Secondary phloem (mg/g)^b			P ^c
	Mean	Range	SD	Mean	Range	SD	P ^c
Petroleum ether–diethyl ether extract	62.31	45.56–84.52	14.03	48.70	31.62–74.63	13.69	<0.001
Diterpenoids
13-Epimanool (1)	1.97	0.71–3.47	0.96	1.52	0.34–3.09	0.90	<0.001
Larixol (2)	3.84	2.09–6.25	1.27	2.47	1.26–3.47	0.75	<0.001
Larixyl acetate (3)	2.28	0.40–4.00	1.09	1.51	0.34–3.22	0.76	<0.001
13-Epitorulosyl acetate (4)	1.87	0.96–3.24	0.66	1.20	0.49–2.25	0.57	<0.001
Abietic acid (5)	1.86	0.91–3.09	0.67	1.07	0.40–1.95	0.43	<0.001
Neoabietic acid (6)	0.27	0.12–0.44	0.11	0.24	0.11–0.41	0.11	0.001
Dehydroabietic acid (7)	0.60	0.32–1.03	0.26	0.39	0.20–0.64	0.13	0.001
Isopimaric acid (8)	2.17	1.44–3.31	0.59	1.37	0.62–2.84	0.65	<0.001
Total of the diterpenoids	14.60	8.24–19.78	3.59	9.53	6.00–11.88	1.99	<0.001

n = 12
^aBased on the dry weight of the rhytidome
^bBased on the dry weight of the secondary phloem
^c P values resulted from analysis of the differences in the ether extract and diterpenoid contents between the rhytidome and secondary phloem in the branch bark of each individual tree using the Wilcoxon signed rank test